Summary Sieve-tube plastids of Aristolochia (5 species investigated) contain several starch grains and always one large crystalloid. In Asarum (3 species investigated) starch has not been found in the sieve-tubes. Their plastids contain several cuneate crystalloids that are sometimes arranged around an invisible centre. Asarum sieve-tube plastids look almost like typical plastids of monocotyledon sieve-tubes.-Crystalloids of Aristolochia and of Asarum sieve-tube plastids are composed of 50–60 Å subunits in straight and parallel order as crystalloids in monocotyledon sieve-tube plastids are.The results of the investigations of the fine structure are discussed in relation to the position of the Aristolochiaceae in the system of angiosperms.